We have a machine that we upgraded today. After the upgrade, I am getting connection refused on any emails from a remote host that uses an alias. If I send email directly to a user on the upgraded host, from a remote host, that works fine. If I send email to an alias on the ugraded machine, from the upgraded machine, that also works. What doesn't work is if I sent email from a remote machine to an alias on the upgraded machine, I get a 'connection refused'.
You might want to turn up the logging levels on a couple of the machine's your testing with and see what you can learn. Perhaps the following link will get you going in the right direction.
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By the time I got into work, the problem was resolved. The entire /etc/mail directory was restored and that fixed it. I don't really see any diffrence in the contents of /etc/mail, but I do see a difference in permissions:
drwx------ 3 root root 1.0K Feb 19 17:01 mail/
drwxr-xr-x 3 root root 1.0K Feb 20 06:46 mail.20130220.RSF/
Thanks for posting back the solution.